Bottomland Hardwood

habitat photo
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Aerial view of floodplain forest along the Wolf River below Navarino Wildlife Area, Shawano County. Photo by Eric Epstein.

Habitat Description

Habitat Crosswalk


Bottomland hardwoods occur along rivers and streams, mainly in the southern half of the state but also at scattered locations in the north.  The largest tracts are found along the Mississippi and lower Wisconsin Rivers, with significant stands also occurring on the Chippewa, Black, Yellow, Baraboo, Wolf, Sugar, Rock, St. Croix, and lower Peshtigo.  Small stands are found along many rivers and streams (Mossman 1988).  Important canopy species include silver maple, green ash, river birch, swamp white oak, red maple, black willow, cottonwood, and hackberry.  Several tree species more typical of the southern U.S., such as sycamore, honey locust, and Kentucky coffee tree can be found locally among bottomland hardwoods in southern Wisconsin.  American elm, formerly an important canopy species in these forests, has been greatly reduced by Dutch elm disease and now rarely reaches the canopy before succumbing, although young trees are still fairly common.  Common understory species, often occurring in a patchy distribution, are wood nettle, jewelweed, sedges and grasses, green dragon, cardinal flower, and green-headed coneflower.  Vines such as woodbine, poison ivy, wild grape, moonseed, and wild cucumber can be prominent.  Canopy openings may be invaded by thickets of native shrubs such as prickly ash and dogwoods, while sloughs and the margins of oxbow ponds often have the water-loving buttonbush.

A related forest type, swamp hardwood (lacustrine swamp or NHI’s southern hardwood swamp), occurs in isolated lowland basins and sometimes lakeshores.  It often is dominated by red maple and ashes, including black ash, with a prevalence in the understory of ferns and shrubs such as dogwoods, nannyberry, and alder.  This forest type is influenced by standing water (seasonally high water table or inundation during spring runoff or major precipitation events) rather than by flood waters that flow through the stand, a hydrologic difference that leads to growth rates and understory composition distinct from bottomland hardwoods (WDNR 2011).  Some streamside tracts have characteristics that are intermediate between these two major forest types (e.g., dominated by swamp white oak and black ash).

Bottomland hardwood forests typically vary in structure and composition at both local and landscape scales due to the effects of variations in topography, soils and hydrology, and natural processes such as beaver activity and windthrow.  Indeed, many such forests are naturally interrupted by sloughs, ponds, shrub swamps and meadows, and by even drier barrens habitats on sandy terraces, but these habitats are often interconnected in extensive mosaics with gradual, natural ecotones.  More so than upland forests, which today tend to be separated from other habitats by hard, artificial edges, bottomland hardwoods may best be seen as a dynamic component of a larger ecosystem.

In general, wet floodplain sites tend to be dominated by silver maple and green ash.  River birch is sometimes common, especially along the lower Wisconsin and Mississippi rivers and the lower reaches of their tributaries.  Active or deadwater sloughs, stream and river channels that often interrupt floodplain forest cover, are overhung to varying extents by the spreading, hanging canopies of silver maple or cottonwood, and formerly American elm.  Higher, wet-mesic sites have these same species but with increased prevalence of swamp white oak, hackberry, white ash, basswood, and (on higher, sandier terraces) black oak and sometimes white pine.  With fewer openings than wet forests, and more prevalent shrub and sapling layers, wet-mesic forest structure tends to resemble that of upland forest.  In areas of former glacial lakes (e.g., Central Sand Plains and Hills), or along the conduits that drained them (e.g., Lower Chippewa and especially Lower Wisconsin), terraces can be sandy, and xeric for much of the year, adding an additional component of “river barrens” (Sample and Mossman 1997) to the floodplain mosaic, characterized by sparse cover of grasses and forbs, open sand, river birch, green ash, red cedar and black oak.  On some expansive terraces with deep, sterile sands, these species are joined by jack pine and sand prairie herbs.

Newly exposed wet soil tends to be pioneered by thickets of sandbar willow shrubs, black willow, river birch and cottonwood, while thicker or higher, somewhat drier deposits on higher terraces often are invaded by river birch, cottonwood, green ash, oaks and prickly-ash.

Figure 1 illustrates typical physiographic relationships among floodplain habitat types.  At any point along a river, the cross-section may exhibit only some of these features, and in some cases rises immediately to upland cliffs or bluffs.

Figure 1. Typical Cross-section of a Bottomland System (from USDA Forest Service 2008), with characteristic Wisconsin floodplain communities.

Historical and Present-day Context and Distribution

Acreage estimates for presettlement and current bottomland hardwood forests vary considerably according to definitions and methods, although all accounts acknowledge its limited distribution and extent, and further, that swamp hardwoods were and remain much less common than bottomland (floodplain) hardwoods.  Curtis (1959) considered that both types comprised 420,000 acres (1.2% of the Wisconsin land area) prior to Euro-American settlement, and that 290,000 acres (69% of presettlement) was estimated by the Land Economic Inventory during 1933-47.  Mossman and Matthiae (1988) stated that only 32,000 acres (8%) remained in moderate to high quality stands.  Recent estimates from Forest Inventory and Analysis (FIA) data from Wisconsin estimate 464,000 acres of floodplain and swamp hardwood forest statewide, but based on what may be an inadequate sample.  The best statewide estimate is probably that of the Wisconsin Wetlands Inventory: 686,000 acres. (Calvin Lawrence WDNR, pers. comm., 3 May 2011).

Although these forests have fared better than other native communities in Wisconsin due to the difficulty of converting them to other land uses, they still have declined greatly in extent since European settlement.  Remaining tracts have been fragmented and degraded by a variety of human activities including high-grade logging, grazing, ditching, clearing for agriculture, invasion by exotic species, and alteration of flood regimes on many streams and rivers due to dam construction, road construction, channelization, wetland drainage, and urban development (Mossman 1988; WDNR 2011).  Some new floodplain forest has developed since European settlement due to succession on meadow and oak savanna sites (sometimes after a period of cultivation), facilitated by reduced flooding and the absence of fire.

The largest tracts of bottomland hardwoods occur in southern Wisconsin in the Western Coulee and Ridges Ecological Landscape, with significant acreages also found in the Central Sand Plains and Southeast Glacial Plains.  Other Ecological Landscapes that contain important sites are: Central Lake Michigan Coastal; Central Sand Hills; Forest Transition; North Central Forest; Northern Lake Michigan Coastal; Northwest Lowlands; Superior Coastal Plain; and Western Prairie (WDNR 2008a).

Hardwood swamps have suffered from drainage, ditching, siltation, excessive nutrient input, and overharvest and remain in much smaller tracts than bottomland hardwoods.  Isolated tracts occur nearly statewide.  Northward and in the Central Sand Plains they often are associated with and grade into coniferous and mixed swamps.

Natural Disturbances and Threats

Bottomland hardwoods and other floodplain habitats are adapted to—and driven by—disturbance.  Periodic flooding, particularly in spring, is the primary natural disturbance that historically has shaped this community.  The frequency, timing, duration, and extent of flooding can alter floodplain topography and influence the species composition and structure of both canopy and understory vegetation layers.  Flooding can cause scouring effects from water, ice, and debris that damage or remove vegetation and expose sand or mud on spits or slough margins, can leave tangles of dead branches and other detritus, and can deposit sediments containing nutrients and organic matter that alter the microtopography of the floodplain.  Floods can also carry in seeds and other propagules of plant species (WDNR 2008a; Mossman 1988; WDNR 2011).  Many bottomland tree species are early-successional and are adapted to exploit the conditions created by periodic floods.  Bottomland hardwoods tend to be fast-growing, utilizing the relatively high levels of nutrients and moisture supplied in floodplains to maintain rapid growth.  Several species take advantage of, or require, bare soil for seed germination.  For example, silver maple requires a fresh deposit of silty soil as a seedbed.  Flooding affects stand structure by influencing the survivorship of seedlings based on their location in the floodplain.  Seedlings that become established on the higher elevations of a floodplain are more likely to survive the effects of subsequent floods than those established on the lower elevations.  At lower elevations this can result in stands with widely spaced canopy trees and sparse woody understory, with a dense groundlayer of wood nettles and a few other herbs that respond quickly to floodwater recession.  Higher sites typically exhibit more regeneration, a denser shrub layer, and a more diverse herbaceous layer.

Other natural disturbances shape the structure, composition and landscape pattern of bottomland hardwoods as well.  Most of the common tree species are shallow-rooted and subject to throw by windstorms, which can be funneled along river valleys.  Canopy disturbance can range from minor to severe, depending on the wind event.  Beaver can fell saplings and even large canopy trees, and create impoundments that flood out forest tracts, which may, after drainage, persist as open meadows.  Historically, fire was necessary to maintain floodplain savannas (typically dominated by swamp white oak or bur oak), black oak-jack pine barrens on more xeric terraces, and some marshes, meadows and shrub swamps that characteristically interrupt forest cover, especially near uplands.  In Wisconsin, fire has been frequent both historically and currently in portions of the lower Chippewa, lower Black, and lower Wisconsin rivers, and formerly occurred on the Mississippi.  Stands on sand and gravel may be predisposed to fire after periods of seasonal drought (Curtis 1959; WDNR 2008b).

Threats to bottomland hardwoods include altered hydrology, exotic species, excessive herbivory, and conversion, fragmentation, and degradation of the habitat due to a variety of human activities.  Threats to hydrology are of primary significance as the hydrologic regime is such a defining feature of this community, affecting many of its physical characteristics.  Dam and impoundment construction on many rivers and streams has affected the timing, frequency, duration, and magnitude of flood events, thus altering the major natural disturbance to which bottomland hardwoods are adapted, with consequences for species composition and structure, and future successional pathways (WDNR 2008a).  This type of flow regulation often is characterized by flooding that is less extreme but more frequent than that experienced under a natural flood regime, resulting in a lack of erosion and deposition that creates ‘new land’ where early-successional species can thrive, or more frequent inundation of stands that does not allow trees to become established.  A lack of severe flooding may be leading to community succession based not on flood tolerance, as is the case under more natural flood regimes, but on shade tolerance.  For example, reduced flooding on the lower Wisconsin River has resulted in the loss of early-successional or “pioneer” species such as cottonwood and black willow and a shift to a later-successional stage dominated by silver maple and Central Hardwoods species such as bitternut hickory and hackberry which were not historically dominant (Hale et al. 2008).  Artificially high water levels created by impoundment in Upper Mississippi River floodplain forests have replaced large forest tracts with open water, while in remaining stands eliminated less flood-tolerant species such as green ash, swamp white oak, and hackberry and creating near monocultures of the more flood-tolerant silver maple, often in even-aged stands (Knutson and Klaas 1998).

Exotic diseases and insects also have affected species composition and structure.  Dutch elm disease has virtually eliminated American elm as a canopy dominant, opening up the canopy in many bottomland hardwood forests and leaving gaps that in some cases have yet to fill in with trees (Mossman 1988; WDNR 2008a).  Elms were abundant, sometimes very large, and had distinctive limb architecture, so their loss has affected stand structure.  An introduced insect, the emerald ash borer, is expanding its range in Wisconsin and threatens the ash component of the bottomland hardwood community.  Another introduced insect, the gypsy moth, threatens oaks and other species (WDNR 2008a).

The exotic, highly invasive reed canary grass is a major threat to bottomland hardwoods, spreading quickly after a disturbance that opens the canopy, such as timber harvest, windthrow, or disease.  It can quickly dominate the ground layer and impede tree regeneration.  Other problematic invasive plants are moneywort, creeping Charlie, common and glossy buckthorns (on wet-mesic sites) and, potentially, common reed (Phragmites).

Bottomland hardwoods have been reduced in extent by clearing for agriculture and development or conversion to other habitat types (e.g., conversion to marsh or shrub wetlands due to dike construction).  Existing tracts can be degraded by neighboring land uses (e.g., altering the flood regime, pollution or sedimentation from adjacent agriculture or construction) and unsustainable logging and grazing.  Excessive herbivory by white-tailed deer or domestic livestock can alter the composition of understory layers, impede tree regeneration, and contribute to the spread of invasive species such as prickly ash and buckthorn.  Unsustainable forest management practices can fragment habitat, alter species composition, and facilitate the spread of invasive plants (WDNR 2008a; Hale et al. 2008).

While Wisconsin’s climate is changing, displaying increases in seasonal temperatures and average annual precipitation over the past 60 years (WICCI 2011), specific effects of these changes on bottomland communities are difficult to predict.  However, the increased probability of more extreme disturbance events, such as heavy downpours that lead to rapid, severe, more frequent or midsummer flooding (Easterling et al. 2000, Jentsch et al. 2007) may have negative consequences for floodplain systems.

A useful definition of a “healthy” or “high quality” forest stand or ecosystem is more elusive for bottomland hardwoods than for most other forest types, considering the uncertainties associated with the long-term effects of important drivers such as: climate change; exotic invasive plants, insect pests and diseases (which have and may continue to eliminate major tree species from the canopy); watershed inputs; and dams (even those that have been in place for many decades).  How will a system dependent on flood regimes—and to a lesser extent on fire, windstorm and beaver activity—respond to this complex of changing influences, most of which are beyond the pale of local forest and habitat managers?  The future of bottomland hardwoods and its associated communities will be best served by a conservative and integrated approach to management that considers the floodplain system and its watershed as a whole, with a long-term perspective that guides monitoring, experimentation, and research.  High quality stands thus include those with intact native flora and fauna, but also those with resilience afforded by size, connectivity, natural ecotones with associated floodplain and upland communities, and limited hydrological disturbance within watersheds and from upriver dams.

Related WBCI Habitats: Shrub-carr, Alder Thicket, Oak Barrens, Oak Opening.

Overall Importance of Habitat for Birds

Bottomland hardwoods in Wisconsin comprise some of the largest remaining forested tracts south of the Tension Zone and support a rich and diverse avifauna (Mossman 1988; Knutson et al. 1996; Miller et al. 2004).  Knutson et al. (1996) reported that species richness and relative abundance of birds in Upper Mississippi River floodplain forests were almost twice as high as those in adjacent upland forests.  Common breeding species are Great Crested Flycatcher, Eastern Wood-Pewee, Song Sparrow, House Wren, American Robin, American Redstart, Baltimore Oriole, Downy Woodpecker, Blue-gray Gnatcatcher, and Blue Jay.  Other less common but characteristic bottomland hardwood species, more abundant in this habitat than in most or all other habitat types in Wisconsin, include Yellow-crowned Night-Heron, Wood Duck, Red-shouldered Hawk, Barred Owl, Yellow-bellied Sapsucker, Tufted Titmouse, Brown Creeper, Warbling Vireo, Prothonotary Warbler, and Kentucky Warbler.  Large tracts of bottomland hardwoods provide important habitat for area-sensitive species, including Red-shouldered Hawk, Acadian Flycatcher, Wood Thrush, Cerulean Warbler, and Kentucky Warbler, particularly where they occur adjacent to extensive tracts of upland forest.  The bottomlands of the Sugar River in far southern Wisconsin where sycamores and other southern tree species extend northward into the state have been an important breeding site for Yellow-throated Warbler (Hansen 2006), an uncommon southern species that also occurs in several upland oak-pine sites in the state.  Although the species has been absent from this area in recent years, it may occur again if climate change causes further northward range shifts of southern tree species into Wisconsin.

The bottomland hardwood avifauna has various distinctive features.  Fish-eating birds, including herons, egrets, Bald Eagle, Osprey, Belted Kingfisher, and Hooded Merganser are well represented.  The large number of dead or dying trees killed by flooding or disease supports a high proportion of cavity nesters, bark gleaners, and wood drillers, including Great Crested Flycatcher, Brown Creeper, Prothonotary Warbler, and 7 of Wisconsin’s 8 breeding woodpeckers.  Red-headed Woodpecker, a species more typically associated with savannas, nested frequently in dead elms in the bottomlands.  Although no longer as common in this habitat with the collapse of many dead elms, mortality of ashes with invasion by emerald ash borer may make this species prevalent in the bottomlands once again.  Bottomland hardwoods also host many cliff-nesting species such as Bank, Barn, Cliff, and Northern Rough-winged Swallows and Belted Kingfisher due to their proximity to cliffs, cutbanks, and bridges (Mossman 1988).  Their location along major river corridors also makes bottomland hardwoods critical habitat for migrating birds, particularly landbirds, in both fall and spring.  Bottomland hardwoods are the main migratory habitat for Rusty Blackbird, the only Wisconsin priority species identified solely on the basis of its presence during migration.

WBCI Priority Bird Species.  Species in boldface are currently proposed as Focal species for southern Wisconsin forests.

Species name Status Habitat and/or Special Habitat Features
Mallard bMwf Generalized, more near urban and open habitats.
Hooded Merganser BMwF Deep sloughs; nest in snags.
Ruffed Grouse bwf Now rare; wet-mesic; where patches of both early-successional and older forest.
Great Egret bMF In extensive floodplain systems of forest-marsh-river-slough-pool; colonies rare, nest with GTBH, may travel far to forage; more widespread before and after nesting season.
Yellow-crowned Night-Heron bmf In extensive floodplain systems of forest-marsh-river-slough-pool.
Bald Eagle BMWF Nest in supercanopy trees; forage on fish in main and side channels, feeder streams, also on upland carrion in winter.
Red-shouldered Hawk BMwF In extensive mature floodplain forest tracts with small pools, sloughs; nest in large trees, sometimes in adjacent upland forest.
American Woodcock bmF In early-successional tracts, large openings, and shrub-carr.
Black-billed Cuckoo bM Mostly migrant; breeds in some shrub-carr, woods edge.
Yellow-billed Cuckoo BM Common but erratic in most floodplain forest tracts >40 acres.
Whip-poor-will bm Disappearing; occurs in wet-mesic sites, more commonly in adjacent barrens or oak woodland.
Chimney Swift bmF Nest in chimneys, occasionally in large “chimney” snags; commonly forage over and within floodplain forest of all types.
Belted Kingfisher BMwF Nest in exposed riverbanks and cliffs, adjacent upland cliffs and cutbanks; forage extensively in channels and sloughs.
Red-headed Woodpecker bmwf Where barkless snags occur due to flooding or disease; also in open-canopy patches.
Yellow-bellied Sapsucker BMw Widespread, especially associated with river birch; nests in cavities in snags and live trees.
Northern Flicker BMw Scattered, where snags, open or semi-open canopy, open herbaceous understory, often using both floodplain forest and adjacent uplands.
Acadian Flycatcher bm Occasional along smaller rivers or at boundary of extensive upland and floodplain forest (especially bases of steep slopes).
Willow Flycatcher bm Shrub carr, most common where least forested.
Least Flycatcher bM Scattered, often in colony-like breeding groups, in both interior and edge.
Yellow-throated Vireo BM Common and widespread in mature forest tracts >40 acres.
Warbling Vireo BM Mostly in forest edge, scattered mature trees, edges of forest sloughs, especially where canopies are spreading or overhanging.
N. Rough-winged Swallow BM Nest in exposed riverbanks and cliffs, adjacent upland cliffs and cutbanks, artificial structures, tip-ups; forage extensively over forest, channels and sloughs; usually nest singly or in small groups, or in Bank Swallow colonies.
Bank Swallow BM Nest in colonies in floodplain and upland cutbanks and cliffs, sand pits.
Veery BM Wet-mesic forest where there are dense shrubs or rank, tall, diverse forest herb layer.
Wood Thrush BM Wet-mesic and adjacent upland forest in tracts >40 acres; territories often include patches of tall semi-dense saplings.
Brown Thrasher bm Sandy open areas, adjacent open upland grassland and barrens.
Blue-winged Warbler BM Wet-mesic, shrubby forest openings, edges and upland-floodplain boundary.
Golden-winged Warbler bm Rare; in same sites as Blue-winged Warbler.
Chestnut-sided Warbler bM Rare breeder, in shrubby forest openings, edges and upland-floodplain boundary.
Yellow-throated Warbler bm Formerly among sycamores along Sugar River and possibly lower Wisconsin and Mississippi rivers; only recent records are from upland oak with supercanopy white pine.
Cerulean Warbler BM Mature forest with diverse canopy species and structure, much more likely where adjacent to extensive upland forest; entire patch must be >240 acres.
Prothonotary Warbler BM Wet forest with sloughs, low snags; more abundant and generalized (less limited to sloughs) near Wis/Miss River confluence.
Louisiana Waterthrush bm Territories always include suitable adjacent upland or floodplain-forest boundary, where stream gorges or springs enter floodplain.
Kentucky Warbler BM Wet-mesic forest and floodplain-upland boundary, where suitable upland-floodplain forest tract >240 acres; often in dense or semi-dense shrubby openings of natural or logging origin.
Mourning Warbler bm Similar to Kentucky Warbler, but more common, less restricted to large tracts, extends into larger open-canopy patches
Common Yellowthroat BM In large forest openings, along edges of rivers, large sloughs and shrub-carr and marsh, more so in wet than wet-mesic sites.
Swamp Sparrow bm Shrub carr, adjacent forest edge, and larger patches of shrub adjacent to large open sloughs.
Rose-breasted Grosbeak BM Common and widespread in forest interior or edge; prefers open or semi-open canopy with moderate sapling/shrub growth.
Rusty Blackbird mwf Declining migrant, much more common in floodplain forest than elsewhere; forages in small vernal pools and wet leaf litter.


The Upper Mississippi River and Great Lakes Joint Venture (UMRGLJV) 2007 Implementation Plan assigns Wisconsin a habitat objective for the bottomland hardwood cover type (Forested Wetland) of 741 acres for Management and Protection and 247 acres for Restoration and Enhancement, using Prothonotary Warbler as a focal planning species.  These objectives are based on BCR 23 and Wisconsin Prothonotary Warbler population estimates and goals that are extremely low because they are extrapolated from data of the federal BBS, a survey that does not sample bottomland habitats, or relatively rare species like Prothonotary Warbler, well.  Without a reliable estimate of Prothonotary Warbler populations in Wisconsin it is difficult to set a numeric habitat objective for bottomland forests based on this species.  The WBCI Southern Forests committee recommends generating a better estimate, but in the interim, the committee suggests a goal to increase Prothonotary Warbler and other focal species populations within bottomland forests in the Wisconsin portion of BCR 23.

Wisconsin holds the most extensive and high-quality floodplain forests in BCR 23, and the entire Great Lakes region, which affords both opportunity and responsibility.  Certainly, a more complete assessment is needed for this community, its birdlife and the sites, species and management issues most worthy of attention.

Management Recommendations

Landscape-level Recommendations

  1. Maintain and protect existing large (>10,000 acres), contiguous tracts of bottomland hardwoods, particularly where they exist adjacent to large tracts of upland forest (Knutson et al. 2001; LMVJV Forest Resource Conservation Working Group 2007; WDNR 2008a; M.J. Mossman, pers. comm. 2011).
  2. Maintain and protect high-quality examples of bottomland hardwoods of any size, particularly when adjacent to other intact habitats.  Many of the recommendations below for increasing connectivity should be targeted in and around these high-quality remnants.
  3. Wherever possible, use forested corridors to link large blocks and high-quality remnants of forest, and to provide for species migration due to climate change (LMVJV Forest Resource Conservation Working Group 2007).
  4. Where feasible, restore natural hydrologic regimes (Knutson et al. 1996).
  5. Manage bottomland hardwood forests as part of existing natural mosaic of floodplain habitats and ecological gradients from lowlands to uplands (WDNR 2008a).
  6. Use buffers to protect floodplain systems from negative impacts of upstream land uses (e.g., sedimentation, pollution) (WDNR 2008a).
  7. Widen floodplain corridors where feasible, particularly to incorporate natural ecological gradients (e.g., by reforesting reclaimed agricultural land) (Knutson and Klaas 1998).
  8. Maintain or provide a diversity of tree species and age classes across the landscape (Knutson and Klaas 1998; Twedt and Wilson 2007).

Site-level Recommendations

  1. Any management activity in this community must attempt to avoid the introduction and spread of invasive species, particularly reed canary grass, to the understory (WDNR 2008a).
  2. Retain large live trees, cavity trees, snags, and coarse woody debris (Twedt and Wilson 2007).
  3. A variable retention silvicultural approach (Mitchell and Beese 2002) and “wildlife forestry” treatments such as clustered thinning and small (~1-3.5 acres) patch cuts (Twedt and Somershoe 2009; Norris et al. 2009) can be used to create and maintain structural complexity (canopy gaps, understory and sub-canopy layers) and mimic natural disturbance while allowing for regeneration; its effects on forest and bird communities should be monitored.
  4. Monitor levels of white-tailed deer herbivory and lower deer numbers where feasible (WDNR 2008a).

Ecological Opportunities

Ecological Landscape Opportunity Management Recommendations
Central Sand Plains Major All
Southeast Glacial Plains Major All
Western Coulee and Ridges Major All
Central Lake Michigan Coastal Important All
Central Sand Hills Important All
Forest Transition Important All
North Central Forest Important All
Northern Lake Michigan Coastal Important All
Superior Coastal Plain Important All
Western Prairie Important All
Northeast Sands Present 2, 3, 4, 5, 6, 7, 9, 10, 11, 12
Northern Highland Present 2, 3, 4, 5, 6, 7, 9, 10, 11, 12
Northwest Sands Present 2, 3, 4, 5, 6, 7, 9, 10, 11, 12
Southern Lake Michigan Coastal Present 2, 3, 4, 5, 6, 7, 9, 10, 11, 12
Southwest Savanna Present 2, 3, 4, 5, 6, 7, 9, 10, 11, 12

Research Needs


Key Sites

Key Partners

Funding Sources

Information Sources


Contact Information

Steele, Y.  2011.  Bottomland Hardwood Habitat Page.  In Paulios, A. and K. Kreitinger (eds.).  2007-2012.  The Wisconsin All-Bird Conservation Plan, Version 1.0.  Wisconsin Bird Conservation Initiative.  Wisconsin Department of Natural Resources, Madison, WI.

Kreitinger, K., Y. Steele and A. Paulios, editors. 2013.
The Wisconsin All-bird Conservation Plan, Version 2.0. Wisconsin Bird Conservation Initiative.
Wisconsin Department of Natural Resources. Madison, WI.

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